Aspects of this topic are discussed in the following places at Britannica.
...exposure, drying, and other hazards is accomplished in various ways, such as by the formation of the sori in grooves or pockets or by the production of various forms of covers. One is the so-called false indusium, a rolled-over leaf margin under which sporangia form and mature. The true indusium is a separate and unique formation, the structural origins of which are not clear, that constitutes...
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